You can't! What's on paper and the actual gentic makeup of your dog are two
entirely different things. One way to look is on paper. In this way.......
wolf bred to shepherd = 50%wolfdog, wolf bred to shepherd =50% wolfdog these
are what are commonly refered to as an F1 or first generation wolf x dog
cross. The resulting half wolf half dog are 50% because the inherit half
their genes from the wolf and half from the dog.
So the only accurate percentage is the first generation (F1) wolf x dog cross
which is 50%
Now from here out is just gets more complicated. If you were to breed two F1
50% wolf dogs to each other on paper it would add up to 50%, genetically
however it is not so simple, We are dealing with two things that happen to
Chromosomes that mix them up in offspring. They go through two processes
independent assortment and recombination. In independent assortment when germ
cells are made they are split up randomly. Take into consideration that
dogs/wolves each have 78 chromosomes so each parent supplies a total of 39
chromosomes to the embryo. Next you have recombination these chromosomes
actually line up and exchange threads of DNA. So when these two F1 50% mate
the chromosomes will undergo independent assortmaent and recombination,
mixing wolf and dog genes randomly. So to put it bluntly after the 50% there
is no telling which genes are actually passed on to the pup(offspring). Out
of two F1 50% you could get a pup with 37% wolf genes, 45%, 67% or even a
pure wolf or dog though the latter two is almost genetically impossible.
So when all the 2nd gen half wolf predecessors such as Deke, Dinosaur,
Flash, Mingo Zeus etc were born each actually inherited a different
percentage of wolf genes from their 50% parents. I believe if some one was to
phenotypically assess the appearance of the previous mentioned 2nd gen
half-wolves they would see a degree of variance manifested in them.
Okay back to paper the above mentioned half-wolves were bred back to German
Shepherds again, On paper you would have a 25%wolfdog.. 50%halfwolf + German
Shepherd = 25% or what was referred to as the first generation quarterwolf.
Now heres something interesting something to ponder about now that we see how
all these genes are randomly mixed it becomes apparent that you are in for a
long term project when it comes to actually setting type. It takes a few if
not many generations of breeding to set the correct type, size and try to
come to some type of uniform look or appearance. Now quarterwolf was bred to
quarterwolf, 25% to 25% making second and third generation quarterwolves. The
first generation 25% were eventually named ATS of course when we look back at
how the genes randomly combine you can only imagine the RANDOM appearance of
these 25% ATS. The process like I mentioned continued to breed 25% to 25% if
a person goes by a magellen scale you could say that eventually things will
even out and you would either remain at that level or breed it out dependent
on course what a person is purposely selecting in the individual dogs
selected for breeding. BUT even today there is great variance in size, head
shape, body type etc. that exist in the ATS dog breed. 1st generation
quarterwolves are still around even though they are registered as ATS. For
example Ntl Ch. Grizzly of Catania, now here is a question he is out of the
black & tan half-wolf Zeus and a white German Shepherd what can this animal
possibly throw white shepherd? Black & tan ? does his offspring conform true
to type does theirs?
In conclusion I must say the ATS is a purebred dog the days of percentages
are over and highly inaccurate exect in the case of a pure wolf pure dog
mating. In the continuing developement of this breed ATS breeders must focus
of breeding the correct temperament and an agreed upon type as of this day
there seemes to be two types of ATS guess what they are? One favors the wolf
the other the GSD.
Tom Baswell
I guess this means a genetic article on the ATS is going to be in the works
but to summerize quickly. There is no definitive test to assess the amount of
wolf blood in a dog. Wolves and Dogs are nearly gentically indistinguishiable
from one another and domestic dogs genotype differ from wolves by only 0.2%
percent sequence divergence.
Original ideals for article from Nick Federoff
Another related article
Wolf dog percent explained by marbles.
Wolfdogproject-percent